This project continued our study of spatial and temporal variability of shoreline biota in the South and Central Puget Sound Basins. Sampling done in summer 2000 focused on quantifying interannual variation among biota, and testing the stability of the north-south negative trend in species diversity seen in the more extensive 1999 sampling. Preliminary data from our 1999 intertidal surveys of pebble beaches suggested that wave energy gradients along the axis of Puget Sound affect community structure by forcing the removal of fine sediments. In June 2000, we resampled 21 pebble beaches from the original 45 sampled in 1999, to compare data among years. We retained the nested sampling design in order to compare within and among different spatial scales. In each of 7 areas, the biota from three replicate beaches were sampled in the low zone along a 50 meter horizontal transect. The 7 areas consisted of three bays in the southern basin of Puget Sound, and four circulation cells in the central basin. Replicate beaches were selected based on similarity of the geomorphic form, sediment size, slope angle, aspect, wave energy, surface roughness, and pore water chemistry. Data on mean annual water temperature, salinity, air temperature, precipitation, and wind speed and direction wer
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