Gamma-Ray Burst (GRB) afterglows are commonly interpreted as synchrotron
emission from a relativistic blast wave produced by a point explosion in an
ambient medium, plausibly the interstellar medium of galaxies. We calculate the
amplitude of flux fluctuations in the lightcurve of afterglows due to
inhomogeneities in the surrounding medium. Such inhomogeneities are an
inevitable consequence of interstellar turbulence, but could also be generated
by variability and anisotropy in a precursor wind from the GRB progenitor.
Detection of their properties could provide important clues about the
environments of GRB sources. We apply our calculations to GRB990510, where an
rms scatter of 2% was observed for the optical flux fluctuations on the 0.1--2
hour timescale during the first day of the afterglow, consistent with it being
entirely due to photometric noise (Stanek et al. 1999). The resulting upper
limits on the density fluctuations on scales of 20-200 AU around the source of
GRB990510, are lower than the inferred fluctuation amplitude on similar scales
in the Galactic interstellar medium. Hourly monitoring of future optical
afterglows might therefore reveal fractional flux fluctuations at the level of
a few percent.Comment: 18 pages, submitted to Ap