Individual males can adopt alternative mating tactics. The occurrence of satellite males is a common behaviour across anuran taxa (e.g., Lithobates clamitans, Wells, 1977; Anaxyrus cognatus, Krupa, 1989; Dendropsophus ebraccatus, Miyamoto and Cane, 1980; Rhinella crucifer, Forester and Lynken, 1986). Satellite males take peripheral positions to calling males, and adopt alternate mating tactics in an attempt to intercept females that are attracted to calling males (Wells, 2007) to increase their own mating success. Satellite males could have an inexpensive form of mate-locating, avoiding predators, and saving energy (Arak, 1983). Furthermore, this strategy could play an important role in the genetic structure of populations (Lodé and Lesbarrères, 2004)
