A key question of theoretical ecology is which properties of ecosystems favour their stability and help maintaining biodiversity. This qu estion recently reconsid- ered mutualistic systems, generating intense controversy about the role of mutu- alistic interactions and their network architecture. Here we show analytically and verify with simulations that reducing the effective intersp ecific competition and the propagation of perturbations positively influences struct ural stability against envi- ronmental perturbations, enhancing persistence. Notewor thy, mutualism reduces the effective interspecific competition only when the direct interspecific competition is weaker than a critical value. This critical competition i s in almost all cases larger in pollinator networks than in random networks with the same connectance. Highly connected mutualistic networks reduce the propagation of e nvironmental perturba- tions, a mechanism reminiscent of MacArthur’s proposal tha t ecosystem complexity enhances stability. Our analytic framework rationalizes p revious contradictory re- sults, and it gives valuable insight on the complex relation ship between mutualism and biodiversity
