Liphone Nophaseud

Michael Bonkowski

Thomas Ziegler

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80   Accepted by A. Bauer: 29 May 2014; published: 9 Jul. 2014
ZOOTAXA
ISSN 1175-5326  (print edition)
ISSN 1175-5334 (online edition) Copyright © 2014 Magnolia Press
Zootaxa 3835 (1): 080–096  
www.mapress.com/zootaxa/ Article
http://dx.doi.org/10.11646/zootaxa.3835.1.4
http://zoobank.org/urn:lsid:zoobank.org:pub:3C8B1815-2A37-414C-99A9-982E274769AF
A new species of Cyrtodactylus (Squamata: Gekkonidae) from the karst forest of 
northern Laos
NICOLE SCHNEIDER
1, TRUONG QUANG NGUYEN
2,3,8, MINH DUC LE
4,5,6, 
LIPHONE NOPHASEUD
7, MICHAEL BONKOWSKI
2 & THOMAS ZIEGLER
1,2
1AG Zoologischer Garten Köln, Riehler Straße 173, D-50735 Köln, Germany. E-mail: schneider.nicole87@googlemail.com and 
ziegler@koelnerzoo.de 
2Zoological Institute, Department of Terrestrial Ecology, University of Cologne, Zülpicher Strasse 47b, D-50674 Cologne, Germany. 
E-mail: m.bonkowski@uni-koeln.de
3Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam. 
Email: nqt2@yahoo.com
4 Faculty of Environmental Sciences, Hanoi University of Science, Vietnam National University, 334 Nguyen Trai Road, Hanoi, 
Vietnam. E-mail: le.duc.minh@hus.edu.vn
5Centre for Natural Resources and Environmental Studies, Hanoi National University, 19 Le Thanh Tong, Hanoi, Vietnam
6Department of Herpetology, American Museum of Natural History, Central Park West at 79
th Street, New York, New York 10024
7Faculty of Science, National University of Laos, Dong Dok Campus, Vientiane, Lao PDR. E-mail: nophasead2007@yahoo.com
8Corresponding author
Abstract
We describe a new species of the gekkonid genus Cyrtodactylus on the basis of two specimens collected from limestone 
forests of Luang Prabang Province, northern Laos. Morphologically, the new species is distinguishable from its congeners 
by a combination of the following diagnostic characters: maximum SVL 86.1 mm; supralabials 9 or 10; infralabials 7–9; 
dorsal tubercles in 15 or 16 rows at midbody; ventral scale rows 34–36 at midbody; precloacal groove absent; femoral 
scales not distinctly enlarged; precloacal pores absent in females (unknown in males); subdigital lamellae under the fourth 
finger 18 or 19, under the fourth toe 18–20; subcaudals not transversally enlarged; dorsal bands white, 4 or 5 between limb 
insertions plus another one between hind limbs; tail banded. Based on molecular analyses, the new species is clustered in 
the same clade with C. wayakonei and two other species from Luang Prabang and Houaphan provinces. 
Key words: Bent-toed gecko, limestone forest, phylogeny, taxonomy, Luang Prabang Province
Introduction
With 186 recognized species, Cyrtodactylus is one of the most diverse genera of lizards in terms of species richness 
(Uetz & Hosek 2014). Since 2010, 63 species have been described in the genus and most of these new discoveries 
were from Southeast Asian countries (Nguyen et al. 2013; Ziegler et al. 2013; Grismer et al. 2014; Luu et al. 2014; 
Phung et al. 2014; Pauwels & Sumontha 2014; Pauwels et al. 2013, 2014; Schneider et al. 2014; Uetz & Hosek 
2014). A major factor for the high species diversity of bent-toed geckos in Southeast Asia is the presence of a large 
area of tropical rain forest and a number of species complexes. For instance, 10 new species have been described 
within the Cyrtodactylus irregularis species complex from the Indochina region and eight new species were 
discovered in the C. pulchellus species complex from the Thai-Malay Peninsula (Grismer et al. 2012; Sumontha et 
al. 2012; Nguyen et al. 2013; Schneider et al. 2014). In Laos, nine species of Cyrtodactylus are currently 
recognized and seven of them have been recorded from limestone karst forests, namely C. interdigitalis Ulber, 
1993 and C. pageli Schneider, Nguyen, Schmitz, Kingsada, Auer & Ziegler, 2011 from Vientiane Province; C. 
lomyenensis Ngo & Pauwels, 2010, C. roesleri Ziegler, Nazarov, Orlov, Nguyen, Vu, Dang, Dinh & Schmitz, 2010, 
and C. jaegeri Luu, Calame, Bonkowski, Nguyen & Ziegler, 2014 from Khammuane Province; C. wayakonei 
Nguyen, Kingsada, Rösler, Auer & Ziegler, 2010 from Luang Nam Tha Province; and C. teyniei David, Nguyen,  Zootaxa 3835 (1)  © 2014 Magnolia Press  ·  93 NEW CYRTODACTYLUS FROM LAOS
Acknowledgements
We are grateful to S. Wayakone, S. Bounphanmy, B. Praxaysombath (Vientiane) and V. Kanyasone (Luang 
Prabang) for supporting our field research in Laos. Export of collected specimens was done due to the export 
permit Nr. 139/13 signed by the CITES Management Authority of Lao PDR. T. Q. Nguyen thanks A. Teynié 
(Aubière), C. X. Le, T. H. Tran (Hanoi), T. Pagel (Cologne), W. Böhme and D. Rödder (Bonn) for support of his 
research. Thanks to H. T. Duong (Hanoi) for laboratory assistance. We are grateful to A. Bauer (Villanova), S. N. 
Nguyen (Ho Chi Minh City), and O. S. G. Pauwels (Brussels) for reviewing the manuscript. Thanks to E. Sterling 
(New York) and K. Koy (Berkeley) for providing the map. Field survey in Luang Prabang Province was funded by 
the Alexander von Humboldt Foundation for TQN (VIE 1143441) and Cologne Zoo for NS. Research of T. Q. 
Nguyen was funded by the Alexander von Humboldt Foundation (VIE 1143441).
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