We thank the Tsawout and Tseycum First Nations Bands for allowing access to X̱OX̱ DEȽ (Mandarte); everyone who contributed to long-term data collection; NSERC (Canada), the Swiss National Science Foundation, the Research Council of Norway (SFF-III, project 223257) and NTNU for funding; and Bob O’Hara for helpful discussions on analyses.Peer reviewe