Plant Enemy-derived Elicitors Increase the Foliar Tannin Concentration of Onobrychis viciifolia Without a Trade-off to Growth

Background and Aims Molecular experiments suggest that the regulation of the biosynthesis of condensed tannin (CT) is sensitive to the presence of plant enemies. The enemy-specific response of CT concentrations to simulated attacks by pathogenic fungi, bacteria or herbivores was studied in Onobrychis viciifolia grown at four levels of nutrient availability. It was hypothesized that CT concentrations increase in response to an attack, and that constitutive and induced levels of CT are higher at low than at high nutrient availability. Investment in CT was also predicted to be negatively related to plant growth. Methods Recently discovered substances by which plants recognize their opponents (i.e. elicitors) were used to simulate attacks to Onobrychis viciifolia grown at 0·0027, 0·075, 0·67 or 2 mm phosphorus in the nutrient solution. Key Results Relative growth rate and final biomass (P < 0·001) were highest at 0·67 mm of phosphorus. CT concentrations decreased with increasing phosphorus availability, from 94·9 to 69·0 mg g−1 leaf dry weight (P < 0·001). Compared with unscathed plants, sterile mere mechanical wounding reduced tannin concentrations from 83·8 to 69·3 mg g−1 leaf dry weight (P < 0·01). Local CT concentrations were higher when wounded leaves were additionally treated with fungal (+15·9 %), bacterial (+19·6 %) or insect (+31·0 %) elicitors (each elicitor; P < 0·05); however, only the insect elicitor (saliva of the lepidopteron Spodoptera littoralis) induced CT concentrations higher than those of unscathed leaves. Conclusions CT concentrations were inducible in the vicinity of the wound but the level of induction was unrelated to the nutrient status of the plant. There was no evidence of a growth-defence trade-off. The inverse relathionship between CT concentrations and nutrient availability appears to reflect passive growth dilution at high nutrient availability, rather than surplus CT production at low nutrient availabilit

Entwicklung eines regionalen Ausbildungsmodells „Ökolandbau und Vermarktung“ an der Fachhochschule Eberswalde im Rahmen des Bundesförderprogramms Regionen Aktiv

In September 2004 a new B. Sc. study course for Organic Farming and Marketing started at the University of Applied Sciences Eberswalde. Supported by the Federal Ministry of Consumer Protection, Food and Agriculture program “Regionen Aktiv” an innovative education model was established. Focussing on rural development farmers, processors and trading partners were involved in the development of topics for the study programme

The Effects of X-Ray Feedback from AGN on Host Galaxy Evolution

Hydrodynamic simulations of galaxies with active galactic nuclei (AGN) have typically employed feedback that is purely local: i.e., an injection of energy to the immediate neighborhood of the black hole. We perform GADGET-2 simulations of massive elliptical galaxies with an additional feedback component: an observationally calibrated X-ray radiation field which emanates from the black hole and heats gas out to large radii from the galaxy center. We find that including the heating and radiation pressure associated with this X-ray flux in our simulations enhances the effects which are commonly reported from AGN feedback. This new feedback model is twice as effective as traditional feedback at suppressing star formation, produces 3 times less star formation in the last 6 Gyr, and modestly lowers the final BH mass (30%). It is also significantly more effective than an X-ray background in reducing the number of satellite galaxies.Comment: 9 emulateapj pages, 8 figures; accepted to Ap

Deep near-infrared imaging of the HE0450-2958 system

The QSO HE0450-2958 and the companion galaxy with which it is interacting, both ultra luminous in the infrared, have been the subject of much attention in recent years, as the quasar host galaxy remained undetected. This led to various interpretations on QSO and galaxy formation and co-evolution, such as black hole ejection, jet induced star formation, dust obscured galaxy, or normal host below the detection limit. We carried out deep observations in the near-IR in order to solve the puzzle concerning the existence of any host. The object was observed with the ESO VLT and HAWK-I in the near-IR J-band for 8 hours. The images have been processed with the MCS deconvolution method (Magain, Courbin & Sohy, 1998), permitting accurate subtraction of the QSO light from the observations. The compact emission region situated close to the QSO, called the blob, which previously showed only gas emission lines in the optical spectra, is now detected in our near-IR images. Its high brightness implies that stars likely contribute to the near-IR emission. The blob might thus be interpreted as an off-centre, bright and very compact host galaxy, involved in a violent collision with its companion.Comment: 4 pages, 3 figures, accepted for publication in A&

Are Radio AGN Powered by Accretion or Black Hole Spin?

We compare accretion and black hole spin as potential energy sources for outbursts from AGN in brightest cluster galaxies (BCGs). Based on our adopted spin model, we find that the distribution of AGN power estimated from X-ray cavities is consistent with a broad range of both spin parameter and accretion rate. Sufficient quantities of molecular gas are available in most BCGs to power their AGN by accretion alone. However, we find no correlation between AGN power and molecular gas mass over the range of jet power considered here. For a given AGN power, the BCG's gas mass and accretion efficiency, defined as the fraction of the available cold molecular gas that is required to power the AGN, both vary by more than two orders of magnitude. Most of the molecular gas in BCGs is apparently consumed by star formation or is driven out of the nucleus by the AGN before it reaches the nuclear black hole. Bondi accretion from hot atmospheres is generally unable to fuel powerful AGN, unless their black holes are more massive than their bulge luminosities imply. We identify several powerful AGN that reside in relatively gas-poor galaxies, indicating an unusually efficient mode of accretion, or that their AGN are powered by another mechanism. If these systems are powered primarily by black hole spin, rather than by accretion, spin must also be tapped efficiently in some systems, i.e., $P_{\rm jet} > \dot Mc^2$, or their black hole masses must be substantially larger than the values implied by their bulge luminosities. We constrain the (model dependent) accretion rate at the transition from radiatively inefficient to radiatively efficient accretion flows to be a few percent of the Eddington rate, a value that is consistent with other estimates.Comment: Accepted for Publication in ApJ, January 2011, 21 pages, 4 figures; One new data point added to figures; clarifications and minor corrections in response to referee repor

Galaxy Zoo: dust lane early-type galaxies are tracers of recent, gas-rich minor mergers

We present the second of two papers concerning the origin and evolution of local early-type galaxies exhibiting dust features. We use optical and radio data to examine the nature of active galactic nucleus (AGN) activity in these objects, and compare these with a carefully constructed control sample. We find that dust lane early-type galaxies are much more likely to host emission-line AGN than the control sample galaxies. Moreover, there is a strong correlation between radio and emission-line AGN activity in dust lane early-types, but not the control sample. Dust lane early-type galaxies show the same distribution of AGN properties in rich and poor environments, suggesting a similar triggering mechanism. By contrast, this is not the case for early-types with no dust features. These findings strongly suggest that dust lane early-type galaxies are starburst systems formed in gas-rich mergers. Further evidence in support of this scenario is provided by enhanced star formation and black hole accretion rates in these objects. Dust lane early-types therefore represent an evolutionary stage between starbursting and quiescent galaxies. In these objects, the AGN has already been triggered but has not as yet completely destroyed the gas reservoir required for star formation.Comment: 11 pages, 18 figures, 4 tables, MNRAS (Accepted for publication- 2012 January 19

A Powerful AGN Outburst in RBS 797

Utilizing $\sim 50$ ks of Chandra X-ray Observatory imaging, we present an analysis of the intracluster medium (ICM) and cavity system in the galaxy cluster RBS 797. In addition to the two previously known cavities in the cluster core, the new and deeper X-ray image has revealed additional structure associated with the active galactic nucleus (AGN). The surface brightness decrements of the two cavities are unusually large, and are consistent with elongated cavities lying close to our line-of-sight. We estimate a total AGN outburst energy and mean jet power of $\approx 3 - 6 \times 10^{60}$ erg and $\approx 3 - 6 \times 10^{45}$ erg s$^{-1}$, respectively, depending on the assumed geometrical configuration of the cavities. Thus, RBS 797 is apparently among the the most powerful AGN outbursts known in a cluster. The average mass accretion rate needed to power the AGN by accretion alone is $\sim 1 M_{\odot}$ yr$^{-1}$. We show that accretion of cold gas onto the AGN at this level is plausible, but that Bondi accretion of the hot atmosphere is probably not. The BCG harbors an unresolved, non-thermal nuclear X-ray source with a bolometric luminosity of $\approx 2 \times 10^{44}$ erg s$^{-1}$. The nuclear emission is probably associated with a rapidly-accreting, radiatively inefficient accretion flow. We present tentative evidence that star formation in the BCG is being triggered by the radio jets and suggest that the cavities may be driving weak shocks ($M \sim 1.5$) into the ICM, similar to the process in the galaxy cluster MS 0735.6+7421.Comment: Accepted to ApJ; 20 pages, 11 low-resolution figure

The central black hole mass of the high-sigma but low-bulge-luminosity lenticular galaxy NGC 1332

The masses of the most massive supermassive black holes (SMBHs) predicted by the M_BH-sigma and M_BH-luminosity relations appear to be in conflict. Which of the two relations is the more fundamental one remains an open question. NGC 1332 is an excellent example that represents the regime of conflict. It is a massive lenticular galaxy which has a bulge with a high velocity dispersion sigma of ~320 km/s; bulge--disc decomposition suggests that only 44% of the total light comes from the bulge. The M_BH-sigma and the M_BH-luminosity predictions for the central black hole mass of NGC 1332 differ by almost an order of magnitude. We present a stellar dynamical measurement of the SMBH mass using an axisymmetric orbit superposition method. Our SINFONI integral-field unit (IFU) observations of NGC 1332 resolve the SMBH's sphere of influence which has a diameter of ~0.76 arcsec. The sigma inside 0.2 arcsec reaches ~400 km/s. The IFU data allow us to increase the statistical significance of our results by modelling each of the four quadrants separately. We measure a SMBH mass of (1.45 \pm 0.20) x 10^9 M_sun with a bulge mass-to-light ratio of 7.08 \pm 0.39 in the R-band. With this mass, the SMBH of NGC 1332 is offset from the M_BH-luminosity relation by a full order of magnitude but is consistent with the M_BH-sigma relation.Comment: 15 pages, 12 figures, accepted for publication in MNRA

Urea Activation by an External Brønsted Acid: Breaking Self-Association and Tuning Catalytic Performance

In this work, we hypothesize that Bronsted acids can activate urea-based catalysts by diminishing its self-assembly tendency. As a proof of concept, we used the asymmetric Friedel-Crafts alkylation of indoles with nitroalkenes as a benchmark reaction. The resulting 3-substituted indole derivatives were obtained with better results due to cooperative effects of the chiral urea and a Bronsted acid additive. Such synergy has been rationalized in terms of disassembly of the supramolecular catalyst aggregates, affording a more acidic and rigid catalytic complex

On the offset of barred galaxies from the black Hole MBH-σ relationship

We use collisionless N-body simulations to determine how the growth of a supermassive black hole (SMBH) influences the nuclear kinematics in both barred and unbarred galaxies. In the presence of a bar, the increase in the velocity dispersion σ (within the effective radius) due to the growth of an SMBH is on average �10%, whereas the increase is only �4% in an unbarred galaxy. In a barred galaxy, the increase results from a combination of three separate factors: (1) orientation and inclination effects; (2) angular momentum transport by the bar that results in an increase in the central mass density; and (3) an increase in the vertical and radial velocity anisotropy of stars in the vicinity of the SMBH. In contrast, the growth of the SMBH in an unbarred galaxy causes the velocity distribution in the inner part of the nucleus to become less radially anisotropic. The increase in σ following the growth of the SMBH is insensitive to a variation of a factor of 10 in the final mass of the SMBH, showing that it is the growth process rather than the actual SMBH mass that alters bar evolution in a way that increases σ.We argue that using an axisymmetric stellar dynamical modeling code to measure SMBH masses in barred galaxies could result in a slight overestimate of the derived MBH, especially if a constant M/L ratio is assumed. We conclude that the growth of a black hole in the presence of a bar could result in an increase in σ that is roughly 4%–8% larger than the increase that occurs in an axisymmetric system. While the increase in σ due to SMBH growth in a barred galaxy might partially account for the claimed offset of barred galaxies and pseudo bulges from the MBH–σ relation obtained for elliptical galaxies and classical bulges in unbarred galaxies, it is inadequate to account for all of the offset