The identity of belowground herbivores, not herbivore diversity, mediates impacts on plant productivity

Across many ecosystems, increases in species biodiversity generally results in greater resource acquisition by consumers. Few studies examining the impacts of consumer diversity on resource capture have focused on terrestrial herbivores, however, especially taxa that feed belowground. Here we conducted field mesocosm experiments to examine the effects of variation in species richness and composition within a community of wireworm herbivores on wheat plant productivity. Our experiments involved wireworm communities consisting of between one and three species, with all possible combinations of species represented. We found that the presence of wireworms reduced plant biomass and seed viability, but wireworm species richness did not impact these plant metrics. Species identity effects were strong, as two species, Limonius californicus and Selatosomus pruininus, had significantly stronger impacts on plants compared to L. infuscatus. Communities with either of the two most impactful species consistently had the greatest impact on wheat plants. The effects of wireworms were thus strongly dependent on the particular species present rather than the overall diversity of the wireworm community. More broadly, our study supports the general finding that the identity of particular consumer species within communities often has greater impacts on ecosystem functioning than species richness

The PROIEL treebank family: a standard for early attestations of Indo-European languages

This article describes a family of dependency treebanks of early attestations of Indo-European languages originating in the parallel treebank built by the members of the project Pragmatic Resources in Old Indo-European Languages (PROIEL). The treebanks all share a set of open-source software tools, including a web annotation interface, and a set of annotation schemes and guidelines developed especially for the project languages. The treebanks use an enriched dependency grammar scheme complemented by detailed morphological tags, which have proved sufficient to give detailed descriptions of these richly inflected languages, and which have been easy to adapt to new languages. We describe the tools and annotation schemes and discuss some challenges posed by the various languages that have been annotated. We also discuss problems with tokenisation, sentence division and lemmatisation, commonly encountered in ancient and mediaeval texts, and challenges associated with low levels of standardisation and ongoing morphological and syntactic change

Jane Austen free indirect style, gender and interiority in literary fiction

In this chapter I suggest that Jane Austen’s use of free indirect style has a far-reaching legacy in terms of establishing the form as central to a sense of literariness in prose fiction. More particularly, I argue that Austen’s use of language metarepresents the thoughts of female characters as a dynamic process of understanding themselves and their worlds. This coincides with a more general perception, construction and performance of ‘feminine’ thought and language use as hesitant, equivocal and spontaneous. I explore the influence of Austen’s style with close analysis and comparison of passages of interiority in Austen’s Mansfield Park, Katherine Mansfield’s short story ‘Millie’ and Monica Ali’s novel Brick Lane.N/