The macroecology of chemical communication in lizards: do climatic factors drive the evolution of signalling glands?

Chemical communication plays a pivotal role in shaping sexual and ecological interactions among animals. In lizards, fundamental mechanisms of sexual selection such as female mate choice have rarely been shown to be influenced by quantitative phenotypic traits (e.g., ornaments), while chemical signals have been found to potentially influence multiple forms of sexual and social interactions, including mate choice and territoriality. Chemical signals in lizards are secreted by glands primarily located on the edge of the cloacae (precloacal glands, PG) and thighs (femoral glands), and whose interspecific and interclade number ranges from 0 to >100. However, elucidating the factors underlying the evolution of such remarkable variation remains an elusive endeavour. Competing hypotheses suggest a dominant role for phylogenetic conservatism (i.e., species within clades share similar numbers of glands) or for natural selection (i.e., their adaptive diversification results in deviating numbers of glands from ancestors). Using the prolific Liolaemus lizard radiation from South America (where precloacal glands vary from 0-14), we present one of the largest-scale tests of both hypotheses to date. Based on climatic and phylogenetic modelling, we show a clear role for both phylogenetic inertia and adaptation underlying gland variation: (i) solar radiation, net primary productivity, topographic heterogeneity and precipitation range have a significant effect on number of PG variation, (ii) humid and cold environments tend to concentrate species with a higher number of glands, (iii) there is a strong phylogenetic signal that tends to conserve the number of PG within clades. Collectively, our study confirms that the inertia of niche conservatism can be broken down by the need of species facing different selection regimes to adjust their glands to suit the demands of their specific environments

Constructing a WISE High Resolution Galaxy Atlas

After eight months of continuous observations, the Wide-field Infrared Survey Explorer (WISE) mapped the entire sky at 3.4 {\mu}m, 4.6 {\mu}m, 12 {\mu}m and 22 {\mu}m. We have begun a dedicated WISE High Resolution Galaxy Atlas (WHRGA) project to fully characterize large, nearby galaxies and produce a legacy image atlas and source catalogue. Here we summarize the deconvolution technique used to significantly improve the spatial resolution of WISE imaging, specifically designed to study the internal anatomy of nearby galaxies. As a case study, we present results for the galaxy NGC 1566, comparing the WISE super-resolution image processing to that of Spitzer, GALEX and ground-based imaging. The is the first paper in a two part series; results for a much larger sample of nearby galaxies is presented in the second paper.Comment: Published in the AJ (2012, AJ, 144, 68

Characterizing the Mid-infrared Extragalactic Sky with WISE and SDSS

The Wide-field Infrared Survey Explorer (WISE) has completed its all-sky survey in four channels at 3.4-22 μm, detecting hundreds of millions of objects. We merge the WISE mid-infrared data with optical data from the Sloan Digital Sky Survey (SDSS) and provide a phenomenological characterization of WISE extragalactic sources. WISE is most sensitive at 3.4 μm (W1) and least sensitive at 22 μm (W4). The W1 band probes massive early-type galaxies out to z ≳ 1. This is more distant than SDSS identified early-type galaxies, consistent with the fact that 28% of 3.4 μm sources have faint or no r-band counterparts (r > 22.2). In contrast, 92%-95% of 12 μm and 22 μm sources have SDSS optical counterparts with r ≤ 22.2. WISE 3.4 μm detects 89.8% of the entire SDSS QSO catalog at S/N_(W1) >7σ, but only 18.9% at 22 μm with S/N_(W4) > 5σ. We show that WISE colors alone are effective in isolating stars (or local early-type galaxies), star-forming galaxies, and strong active galactic nuclei (AGNs)/QSOs at z ≾ 3. We highlight three major applications of WISE colors: (1) Selection of strong AGNs/QSOs at z ≤ 3 using W1 – W2 > 0.8 and W2 0.8, W2 6 (Vega) colors can be used to identify type-2 AGN candidates. The fraction of these type-2 AGN candidates is one-third of all WISE color-selected AGNs. (3) Selection of ultraluminous infrared galaxies (ULIRGs) at z ~ 2 with extremely red colors, r – W4 > 14 or well-detected 22 μm sources lacking detections in the 3.4 and 4.6 μm bands. The surface density of z ~ 2 ULIRG candidates selected with r – W4 > 14 is 0.9 ± 0.07 deg^(–2) at S/N_(W4) ≥ 5 (the corresponding, lowest flux density of 2.5 mJy), which is consistent with that inferred from smaller area Spitzer surveys. Optical spectroscopy of a small number of these high-redshift ULIRG candidates confirms our selection, and reveals a possible trend that optically fainter or r – W4 redder candidates are at higher redshifts

Mid-Infrared Selection of Active Galactic Nuclei with the Wide-Field Infrared Survey Explorer. II. Properties of WISE-Selected Active Galactic Nuclei in the NDWFS Boötes Field

Stern et al. presented a study of Wide-field Infrared Survey Explorer (WISE) selection of active galactic nuclei (AGNs) in the 2 deg^2 COSMOS field, finding that a simple criterion W1-W2 ≥ 0.8 provides a highly reliable and complete AGN sample for W2 < 15.05, where the W1 and W2 passbands are centered at 3.4 μm and 4.6 μm, respectively. Here we extend this study using the larger 9 deg^2 NOAO Deep Wide-Field Survey Boötes field which also has considerably deeper WISE observations than the COSMOS field, and find that this simple color cut significantly loses reliability at fainter fluxes. We define a modified selection criterion combining the W1–W2 color and the W2 magnitude to provide highly reliable or highly complete AGN samples for fainter WISE sources. In particular, we define a color-magnitude cut that finds 130 ± 4 deg^(–2) AGN candidates for W2 < 17.11 with 90% reliability. Using the extensive UV through mid-IR broadband photometry available in this field, we study the spectral energy distributions of WISE AGN candidates. We find that, as expected, the WISE AGN selection can identify highly obscured AGNs, but that it is biased toward objects where the AGN dominates the bolometric luminosity output. We study the distribution of reddening in the AGN sample and discuss a formalism to account for sample incompleteness based on the step-wise maximum-likelihood method of Efstathiou et al. The resulting dust obscuration distributions depend strongly on AGN luminosity, consistent with the trend expected for a receding torus. At L_AGN ~ 3 × 10^(44) erg s^–1, 29% ± 7% of AGNs are observed as Type 1, while at ~4 × 10^(45) erg s^(–1) the fraction is 64% ± 13%. The distribution of obscuration values suggests that dust in the torus is present as both a diffuse medium and in optically thick clouds

Characterizing the Mid-IR Extragalactic Sky with WISE and SDSS

The Wide-field Infrared Survey Explorer (WISE) has completed its all-sky survey at 3.4-22 micron. We merge the WISE data with optical SDSS data and provide a phenomenological characterization of mid-IR, extragalactic sources. WISE is most sensitive at 3.4micron(W1) and least at 22micron(W4). The W1 band probes massive early-type galaxies out to z\gtrsim1. This is more distant than SDSS identified early-type galaxies, consistent with the fact that 28% of 3.4micron sources have faint or no r-band counterparts (r>22.2). In contrast, 92-95% of 12 and 22micron sources have SDSS optical counterparts with r<22.2. WISE 3.4micron detects 89.8% of the entire SDSS QSO catalog at SNR(W1)>7, but only 18.9% at 22micron with SNR(W4)>5. We show that WISE colors alone are effective in isolating stars (or local early-type galaxies), star-forming galaxies and strong AGN/QSOs at z<3. We highlight three major applications of WISE colors: (1) Selection of strong AGN/QSOs at z0.8 and W2<15.2 criteria, producing a census of this population. The surface density of these strong AGN/QSO candidates is 67.5+-0.14/deg^2. (2) Selection of dust-obscured, type-2 AGN/QSO candidates. We show that WISE W1-W2>0.8, W2<15.2 combined with r-W2>6 (Vega) colors can be used to identify type-2 AGN candidates. The fraction of these type-2 AGN candidates is 1/3rd of all WISE color-selected AGNs. (3) Selection of ULIRGs at z\sim2 with extremely red colors, r-W4>14 or well-detected 22micron sources lacking detections in the 3.4 and 4.6micron bands. The surface density of z~2 r-W4>14 ULIRGs is 0.9+-0.07/deg^2 at SNR(W4)>5 (flux(W4)>=2.5mJy), which is consistent with that inferred from smaller area Spitzer surveys. Optical spectroscopy of a small number of these high-redshift ULIRGs confirms our selection, and reveals a possible trend that optically fainter or r-W4 redder candidates are at higher redshifts.Comment: 41 pages, 20 figures, Accepted for publication by the Astronomical Journa

Macroevolutionary diversification of glands for chemical communication in squamate reptiles

Chemical communication plays a central role in social, sexual and ecological interactions among animals. However, the macroevolutionary diversification of traits responsible for chemical signaling remains fundamentally unknown. Most research investigating evolutionary diversification of glands responsible for the production of chemical signals has focused on arthropods, while its study among vertebrates remains neglected. Using a global-scale dataset covering > 80% (7,904 species) of the living diversity of lizards and snakes (squamates), we investigate rates, trajectories and phylogenetic patterns of diversification of their follicular glands for chemical communication. We observed these glands in 13.66% of species, that their expression has varying phylogenetic signal among lineages, and that the crown squamate ancestor lacked follicular glands, which therefore originated and diversified subsequently during their evolutionary history. Additionally, our findings challenge the longstanding view that within squamates the Iguania are visually oriented while Scleroglossa are chemically-oriented, given that Iguania doubles Scleroglossa in the frequency of glands. Our phylogenetic analyses identified stabilizing selection as the best model describing follicular gland diversification, and revealed high rates of disparity. We provide the first global-scale analysis investigating the diversification of one of the main forms of communication among reptiles, presenting a macroevolutionary angle to questions traditionally explored at microevolutionary scale

Extending the Nearby Galaxy Heritage with WISE: First Results from the WISE Enhanced Resolution Galaxy Atlas

The Wide-field Infrared Survey Explorer (WISE) mapped the entire sky at mid-infrared wavelengths 3.4 μm, 4.6 μm, 12 μm, and 22 μm. The mission was primarily designed to extract point sources, leaving resolved and extended sources, for the most part, unexplored. Accordingly, we have begun a dedicated WISE Enhanced Resolution Galaxy Atlas (WERGA) project to fully characterize large, nearby galaxies and produce a legacy image atlas and source catalog. Here we demonstrate the first results of the WERGA project for a sample of 17 galaxies, chosen to be of large angular size, diverse morphology, and covering a range in color, stellar mass, and star formation. It includes many well-studied galaxies, such as M 51, M 81, M 87, M 83, M 101, and IC 342. Photometry and surface brightness decomposition is carried out after special super-resolution processing, achieving spatial resolutions similar to that of Spitzer Infrared Array Camera. The enhanced resolution method is summarized in the first paper of this two-part series. In this second work, we present WISE, Spitzer, and Galaxy Evolution Explorer (GALEX) photometric and characterization measurements for the sample galaxies, combining the measurements to study the global properties. We derive star formation rates using the polycyclic aromatic hydrocarbon sensitive 12 μm (W3) fluxes, warm-dust sensitive 22 μm (W4) fluxes, and young massive-star sensitive ultraviolet (UV) fluxes. Stellar masses are estimated using the 3.4 μm (W1) and 4.6 μm (W2) measurements that trace the dominant stellar mass content. We highlight and showcase the detailed results of M 83, comparing the WISE/Spitzer results with the Australia Telescope Compact Array H I gas distribution and GALEX UV emission, tracing the evolution from gas to stars. In addition to the enhanced images, WISE's all-sky coverage provides a tremendous advantage over Spitzer for building a complete nearby galaxy catalog, tracing both stellar mass and star formation histories. We discuss the construction of a complete mid-infrared catalog of galaxies and its complementary role of studying the assembly and evolution of galaxies in the local universe

Accounting students and communication apprehension: a study of Spanish and UK students

Accounting is about measuring and communicating. Accounting bodies and employers have expressed opinions, which have been supported by research results, advocating that greater emphasis is placed on the development of communication skills throughout the education and training of accountants. Consequently, an increasing number of accounting programmes now include communication skills as educational objectives or learning outcomes, and have integrated activities into the curriculum specifically to develop these skills. It is important to recognise that certain factors can severely restrict the development of communication skills; a major factor is communication apprehension. Research suggests that the existence of high levels of communication apprehension will make efforts to improve communication skills ineffective. Previous research findings indicate that accounting students have high levels of communication apprehension. This paper compares and contrasts the levels and profiles of communication apprehension exhibited by accounting students at the (UK University) and those at the (ESP University). The levels of communication apprehension are also compared with those of students from other disciplines at the same institutions. The results confirm the high levels of communication apprehension in European accounting students. There are notable differences between the two countries however in certain underlying factors.</p