Functions of agonistic interaction, social dominance and display in a winter population of the great tit, Parus major L.

SIGLEAvailable from British Library Document Supply Centre- DSC:D95948 / BLDSC - British Library Document Supply CentreGBUnited Kingdo

Chemical and Behavioral Ecology of the Coastal Giant Salamander (Dicamptodon tenebrosus)

Intraspecific interactions are an important factor in shaping the population structure of terrestrial salamanders. Both physical interactions and scent-marking are vital components to the establishment of territories and influence dispersion of individuals. However, little is known about the terrestrial interactions of many salamander species, particularly that of the Coastal Giant Salamander (Dicamptodon tenebrosus). In this study, I observed the first evidence for the use chemical cues by terrestrial D. tenerosus in regards to shelter selection. I also observed the first accounts of agonistic and display behavior of terrestrial D. tenebrosus. These results suggest that terrestrial D. tenebrosus utilize chemical cues in their selection of refuge sites, and that they also display aggressive, territorial behavior

The effects of weaponry and mating experience on the level and outcome of agonistic interactions in male field crickets, gryllus bimaculatus (orthoptera: gryllidae)

Abstract A wide variety of factors are predicted to influence the intensity and outcome of agonistic interactions in animals, including the resource holding potential of the opponents and the nature and value of the resource over which the individuals are competing. Field crickets (Orthoptera: Gryllidae) have been used extensively as model organisms with which to study animal contests, but relatively few studies have examined the effect of mandible size or structure, or the level of contact with females on the intensity and outcome of agonistic interactions. To do so was the aim of the present thesis, using Gryllus bimaculatus as the study species. The first finding of this study was that there is a significant degree of sexual dimorphism for anterior components of the anatomy in G. bimaculatus. The mandibles, head and pronotum of male crickets were all relatively larger than those of females. This indicates that these traits may be acted upon by intrasexual selection. In many animal species that show sexual dimorphism, a trade-off in development sees enhanced weapon growth at the expense of testes size, but no such relationship was seen in this species. A comparison of the mandible structure of males that either won or lost at flaring and or wrestling showed that a relatively wider mandible span was a significant predictor of success during mandible displays. It was also found that specific components of tooth structure, namely the length of the incisor and length to distal tip, were significantly associated with victory at the jaw flaring stage. This is the first time that mandible shape has been shown to affect fight outcome in the Gryllidae, and also the first confirmed identification of a visual cue component of fighting behaviour. Despite the effectiveness of their weapons in fighting, body mass is a primary predictor of victory in combat between G. bimaculatus males, with the greater the degree of asymmetry in weight the more likely the heavier fighter will win. However, a study of fighting behaviour between asymmetrically matched opponents found that even males who were out-weighed by 40% were still likely to escalate the fight to grappling. Furthermore, males who were able to fend off their larger opponent in their first clash were significantly more likely to win their overall encounter. This hyper-aggressive response may therefore represent an adaptive mechanism to extreme odds and is worthy of further study. Female contact is known to be a significant promoter of male aggression and fighting enthusiasm, and mate guarding aggression is well documented in G. bimaculatus. A recreation of two contradictory studies, including one which concluded that mating makes males lose fights, highlighted that female contact after spermatophore transfer can overcome the loser effect and cause a male to re-engage with a previously dominant opponent. Fighting behaviour in this species is therefore highly flexible and factors affecting the outcome of contests are complex. There is much scope for further studies on this topic.n/

Female Fighting and Host Competition Among Four Sympatric Species of \u3ci\u3eMelittobia\u3c/i\u3e (Hymenoptera: Eulophidae)

Melittobia is a genus of parasitic wasps well known for high levels of inbreeding and violent male combat. Casual observations of groups of sisters of M. femorata placed with hosts revealed a surprising incidence of body mutilations (broken or missing tarsi, antennae, and wings). Replicated conspecific groups of 1, 2, or 3 females of M. femorata, M. digitata, and M. australica and interspecific groups of M. femorata and M. australica (2:1) were observed over their first 10 days in newly established cultures, and the incidence of mutilation was recorded. In some groups females were dye-fed, allowing us to subsequently chart their individual activity patterns on or near the host based on patterns of their colored fecal droppings. For M. australica and M. digitata, no conspecific females in any group size ever showed mutilation. However, in M. femorata nearly 3/4ths of the females in conspecific groups of two or three acquired body damage beginning about the time of first oviposition on the host. In 4 of 5 replicates of the interspecific groups, M. femorata females killed the female of M. australica. Patterns of dyed fecal droppings that developed over several days showed that individual females in groups of both M. femorata and M. australica increasingly restricted their activities to a small portion of the host. These “micro” territories were non-overlapping and appeared to be actively defended. In contrast, M. digitata females in groups never displayed obvious territoriality or interference. Possible reasons for these differences in female behavior are discussed