343,541 research outputs found

    The Influences of Conventional and Low Density Thinning on Leaf Area, Growth, and Growing Space Relationships of Eastern White Pine (Pinus Strobus L.)

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    Despite the commercial importance and widespread management of eastern white pine {Pinus strobus L.) in the Northeast, surprisingly little is known about the effects of thinning on even-aged stand development. To address this, patterns of leaf area, bole form, volume growth, and growth efficiency - defined as volume increment per unit leaf area - were examined over a 17-year period within a thinning study in central Maine designed to compare the conventional B-line and low density thinning regimes. At the tree-level, many of the effects of thinning were as expected. Heavier, low density thinning resulted in significantly larger and deeper crowns with greater leaf area than equivalent trees in both the B-line and unthinned control treatments. These changes explained higher rates of diameter and volume growth. Thinning did not alter growth efficiency per se. but larger trees had slightly (but significantly) lower growth efficiency than smaller trees. Reconstruction of bole taper - quantified as Girard form class -showed that, surprisingly, B-line thinning produced more tapered butt-logs (first 5-meter) than low density thinning, resulting from a thinning-induced growth response at breast height but not at the top of the butt-log. At the stand-level, an annual record of leaf area index (LAI) attained by litterfall collection showed that leaf area in the control treatment was relatively constant or slightly declined over the study period. Thinning significantly reduced stand leaf area and thus gross volume growth, but the thinned treatments had nearly equal LAIs for the ten years following the initial thinning. This explained the similar gross volume growth rates and growth efficiencies of the thinning treatments. Following the re-entry harvest, B-line leaf area increased until the stands reached crown closure, while the low density treatment continued rates of crown expansion and LAI increase without reaching a peak. Due to greater LAI, B-line gross stand volume growth and growth efficiency were significantly higher during the latter growth period: low density stand growth efficiency was still no different from the control. Growth efficiency of the unthinned stands was found to be positively related to stand density. Results of this study have important implications to managers of eastern white pine. The contention that thinning below B-line stocking has deleterious effects on stand yield was in general not supported. On the contrary, only a minor loss of gross stand volume growth was found by thinning to a low density. In addition, low density trees were larger, faster growing, and had better stem form than comparable B-line trees. Therefore, low density thinning was found to be a viable alternative to conventional management

    Leaf Micromorphology Aids Taxonomic Delineation within the Hypervariable Genus Metrosideros (Myrtaceae) on O\u27ahu

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    Variation in leaf micromorphology can aid delimitation of taxonomically difficult groups. The woody genus Metrosideros (Myrtaceae) dominates Hawai\u27i\u27s native forests and comprises striking, predominantly infraspecific, macromorphological variation, including many forms unrecognized in current taxonomic treatments. On taxonomically rich O\u27ahu, 10 taxa (varieties or unnamed morphotypes of M. polymorpha and other species) occur in a predictable sequence with overlapping ranges from low to high elevation along the numerous leeward ridges of the Ko\u27olau Range. We used scanning electron and light microscopy to examine mature, nonsenescing sun leaves from each of these 10 taxa. Parametric and nonparametric tests were used to compare stomatal complex (SC) traits and those of a newly described secretory structure (SS) across taxa. Correlations among traits and discriminant analyses were also done. Density and length of SCs were inversely related overall, and both measures varied significantly across taxa. Although abaxial SSs were consistently present only in the two highest-elevation glabrous taxa, adaxial SSs were present in all 10 taxa. Length and density of adaxial SSs and density of abaxial SSs varied across taxa, and densities of both SCs and SSs were greatest at high elevation. Combined, the SC and SS traits differentiated 100% of four glabrous varieties and two of three pubescent varieties of M. polymorpha. Variation in leaf micromorphology, including presence and density of a newly described SS, aids delimitation of closely related Hawaiian Metrosideros taxa and may refilect differential local adaptation across a heterogeneous landscape

    Lianas Suppress Seedling Growth and Survival of 14 Tree Species in a Panamanian Tropical Forest

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    Lianas are a common plant growth form in tropical forests, where they compete intensely with trees, decreasing tree recruitment, growth, and survival. If the detrimental effects of lianas vary significantly with tree species identity, as is often assumed, then lianas may influence tree species diversity and community composition. Furthermore, recent studies have shown that liana abundance and biomass are increasing relative to trees in neotropical forests, which will likely magnify the detrimental effects of lianas and may ultimately alter tree species diversity, relative abundances, and community composition. Few studies, however, have tested the responses of multiple tree species to the presence of lianas in robust, well‐replicated experiments. We tested the hypotheses that lianas reduce tree seedling growth and survival, and that the effect of lianas varies with tree species identity. We used a large‐scale liana removal experiment in Central Panama in which we planted 14 replicate seedlings of 14 different tree species that varied in shade tolerance in each of 16 80 × 80 m plots (eight liana‐removal and eight unmanipulated controls; 3136 total seedlings). Over a nearly two‐yr period, we found that tree seedlings survived 75% more, grew 300% taller, and had twice the aboveground biomass in liana‐removal plots than seedlings in control plots, consistent with strong competition between lianas and tree seedlings. There were no significant differences in the response of tree species to liana competition (i.e., there was no species by treatment interaction), indicating that lianas had a similar negative effect on all 14 tree species. Furthermore, the effect of lianas did not vary with tree species shade tolerance classification, suggesting that the liana effect was not solely based on light. Based on these findings, recently observed increases in liana abundance in neotropical forests will substantially reduce tree regeneration, but will not significantly alter tropical tree species diversity, relative abundance, or community composition

    Testing mechanisms of compensatory fitness of dioecy in a cosexual world

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    Questions: All else being equal, populations of dioecious species with a 50:50 sex ratio have only half the effective reproductive population size of bisexual species of equal abundance. Consequently, there is a need to explain how dioecious and bisexual species coexist. Increased mean individual seed mass, fecundity, and population density have all been proposed as attributes of unisexual individuals or populations that may contribute to the persistence or resilience of dioecious species. To date, no studies have compared sympatric dioecious and cosexual species with respect to all three components of fitness. In this study, we sought evidence for these compensatory advantages (higher seed mass, greater seed production per unit basal area, and higher population density) in dioecious species. Location: Five 20–25 ha forest dynamic plots spanning a latitudinal gradient in China, including two temperate, two subtropical, and one tropical forest. Methods: We used a phylogenetically corrected generalized linear modelling approach to assess the phylogenetic dependence and joint evolution of sexual system, seed mass and production, and ecological abundances among 48–333 species and 32,568–136,237 individuals per forest. Results: Across all five forests, we detected no consistent advantage for dioecious relative to sympatric cosexual species with respect to mean individual seed mass, seed production or the density of stems in any size class. Conclusions: Our study suggests that seed traits may provide compensatory mechanisms in some forests, but most often the coexistence of sexual systems cannot be explained by advantages of dioecy related to seed quality and demographic parameters. Future investigations of the factors that promote coexistence may increase our understanding by expanding the search to include attributes such as lifespan and tolerance or resistance to herbivores

    Behavioral Modernity and the Cultural Transmission of Structured Information: The Semantic Axelrod Model

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    Cultural transmission models are coming to the fore in explaining increases in the Paleolithic toolkit richness and diversity. During the later Paleolithic, technologies increase not only in terms of diversity but also in their complexity and interdependence. As Mesoudi and O'Brien (2008) have shown, selection broadly favors social learning of information that is hierarchical and structured, and multiple studies have demonstrated that teaching within a social learning environment can increase fitness. We believe that teaching also provides the scaffolding for transmission of more complex cultural traits. Here, we introduce an extension of the Axelrod (1997} model of cultural differentiation in which traits have prerequisite relationships, and where social learning is dependent upon the ordering of those prerequisites. We examine the resulting structure of cultural repertoires as learning environments range from largely unstructured imitation, to structured teaching of necessary prerequisites, and we find that in combination with individual learning and innovation, high probabilities of teaching prerequisites leads to richer cultural repertoires. Our results point to ways in which we can build more comprehensive explanations of the archaeological record of the Paleolithic as well as other cases of technological change.Comment: 24 pages, 7 figures. Submitted to "Learning Strategies and Cultural Evolution during the Paleolithic", edited by Kenichi Aoki and Alex Mesoudi, and presented at the 79th Annual Meeting of the Society for American Archaeology, Austin TX. Revised 5/14/1

    Evaluating the impact of Hymenoscyphus fraxineus in Trentino (Alps, Northern Italy): first investigations

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    The spread of Hymenoscyphus fraxineus has been causing great concern regarding the survival of European ash (Fraxinus excelsior) throughout Europe since the 1990s. The disease was first recorded in Trentino (southern Alps, Italy) in 2012 and has spread throughout the mountain landscape, where ash trees are scattered in small and isolated stands in different valleys. The status of the disease was checked by monitoring the damage to natural regeneration and adult trees in 90 sites spread over the whole region. The survey confirmed the complete colonization by the pathogen of the whole investigated area, with high levels of damage to both young and adult ash trees. Regeneration (both seedlings and saplings) was observed to be affected by the fungus in 88 plots out of 90. Out of 4486 examined young European ashes, 2261 (50.4%) were affected and 789 (17.6%) were already dead. Ten of the 384 assayed flowering ashes (Fraxinus ornus) showed symptoms on branches and apical stems, similar to those observed for European ash. Isolation and molecular analysis proved the presence of the fungus on both symptomatic European and flowering ashes. The examined 386 adult trees showed different levels of damage, sometimes reaching more than 75% of the crown. Some individual trees (42) growing close to severely damaged trees appeared fully healthy, which suggests the possible existence of some resistant/tolerant individuals in the examined populations

    DNA extraction from museum specimens of parasitic Hymenoptera.

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    At the same time that molecular researchers are improving techniques to extract DNA from museum specimens, this increased demand for access to museum specimens has created tension between the need to preserve specimens for maintaining collections and morphological research and the desire to conduct molecular analyses. To address these concerns, we examined the suitability of non-invasive DNA extraction techniques on three species of parasitic Hymenoptera (Braconidae), and test the effects of body size (parasitoid species), age (time since collection), and DNA concentration from each extract on the probability of amplifying meaningful fragments of two commonly used genetic loci. We found that age was a significant factor for determining the probability of success for sequencing both 28S and COI fragments. While the size of the braconid parasitoids significantly affected the total amount of extracted DNA, neither size nor DNA concentration were significant factors for the amplification of either gene region. We also tested several primer combinations of various lengths, but were unable to amplify fragments longer than ~150 base pairs. These short fragments of 28S and COI were however sufficient for species identification, and for the discovery of within species genetic variation

    The statistical neuroanatomy of frontal networks in the macaque

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    We were interested in gaining insight into the functional properties of frontal networks based upon their anatomical inputs. We took a neuroinformatics approach, carrying out maximum likelihood hierarchical cluster analysis on 25 frontal cortical areas based upon their anatomical connections, with 68 input areas representing exterosensory, chemosensory, motor, limbic, and other frontal inputs. The analysis revealed a set of statistically robust clusters. We used these clusters to divide the frontal areas into 5 groups, including ventral-lateral, ventral-medial, dorsal-medial, dorsal-lateral, and caudal-orbital groups. Each of these groups was defined by a unique set of inputs. This organization provides insight into the differential roles of each group of areas and suggests a gradient by which orbital and ventral-medial areas may be responsible for decision-making processes based on emotion and primary reinforcers, and lateral frontal areas are more involved in integrating affective and rational information into a common framework

    The EPOCH Project: I. Periodic variable stars in the EROS-2 LMC database

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    The EPOCH (EROS-2 periodic variable star classification using machine learning) project aims to detect periodic variable stars in the EROS-2 light curve database. In this paper, we present the first result of the classification of periodic variable stars in the EROS-2 LMC database. To classify these variables, we first built a training set by compiling known variables in the Large Magellanic Cloud area from the OGLE and MACHO surveys. We crossmatched these variables with the EROS-2 sources and extracted 22 variability features from 28 392 light curves of the corresponding EROS-2 sources. We then used the random forest method to classify the EROS-2 sources in the training set. We designed the model to separate not only δ\delta Scuti stars, RR Lyraes, Cepheids, eclipsing binaries, and long-period variables, the superclasses, but also their subclasses, such as RRab, RRc, RRd, and RRe for RR Lyraes, and similarly for the other variable types. The model trained using only the superclasses shows 99% recall and precision, while the model trained on all subclasses shows 87% recall and precision. We applied the trained model to the entire EROS-2 LMC database, which contains about 29 million sources, and found 117 234 periodic variable candidates. Out of these 117 234 periodic variables, 55 285 have not been discovered by either OGLE or MACHO variability studies. This set comprises 1 906 δ\delta Scuti stars, 6 607 RR Lyraes, 638 Cepheids, 178 Type II Cepheids, 34 562 eclipsing binaries, and 11 394 long-period variables. A catalog of these EROS-2 LMC periodic variable stars will be available online at http://stardb.yonsei.ac.kr and at the CDS website (http://vizier.u-strasbg.fr/viz-bin/VizieR).Comment: 18 pages, 20 figures, suggseted language-editing by the A&A editorial office is applie
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