The use of multilayer network analysis in animal behaviour

Network analysis has driven key developments in research on animal behaviour by providing quantitative methods to study the social structures of animal groups and populations. A recent formalism, known as \emph{multilayer network analysis}, has advanced the study of multifaceted networked systems in many disciplines. It offers novel ways to study and quantify animal behaviour as connected 'layers' of interactions. In this article, we review common questions in animal behaviour that can be studied using a multilayer approach, and we link these questions to specific analyses. We outline the types of behavioural data and questions that may be suitable to study using multilayer network analysis. We detail several multilayer methods, which can provide new insights into questions about animal sociality at individual, group, population, and evolutionary levels of organisation. We give examples for how to implement multilayer methods to demonstrate how taking a multilayer approach can alter inferences about social structure and the positions of individuals within such a structure. Finally, we discuss caveats to undertaking multilayer network analysis in the study of animal social networks, and we call attention to methodological challenges for the application of these approaches. Our aim is to instigate the study of new questions about animal sociality using the new toolbox of multilayer network analysis.Comment: Thoroughly revised; title changed slightl

Communities in Networks

We survey some of the concepts, methods, and applications of community detection, which has become an increasingly important area of network science. To help ease newcomers into the field, we provide a guide to available methodology and open problems, and discuss why scientists from diverse backgrounds are interested in these problems. As a running theme, we emphasize the connections of community detection to problems in statistical physics and computational optimization.Comment: survey/review article on community structure in networks; published version is available at http://people.maths.ox.ac.uk/~porterm/papers/comnotices.pd

Hypergraphx: a library for higher-order network analysis

From social to biological systems, many real-world systems are characterized by higher-order, non-dyadic interactions. Such systems are conveniently described by hypergraphs, where hyperedges encode interactions among an arbitrary number of units. Here, we present an open-source python library, hypergraphx (HGX), providing a comprehensive collection of algorithms and functions for the analysis of higher-order networks. These include different ways to convert data across distinct higher-order representations, a large variety of measures of higher-order organization at the local and the mesoscale, statistical filters to sparsify higher-order data, a wide array of static and dynamic generative models, and an implementation of different dynamical processes with higher-order interactions. Our computational framework is general, and allows to analyse hypergraphs with weighted, directed, signed, temporal and multiplex group interactions. We provide visual insights on higher-order data through a variety of different visualization tools. We accompany our code with an extended higher-order data repository, and demonstrate the ability of HGX to analyse real-world systems through a systematic analysis of a social network with higher-order interactions. The library is conceived as an evolving, community-based effort, which will further extend its functionalities over the years. Our software is available at https://github.com/HGX-Team/hypergraph

Inferring monopartite projections of bipartite networks: an entropy-based approach

Bipartite networks are currently regarded as providing a major insight into the organization of many real-world systems, unveiling the mechanisms driving the interactions occurring between distinct groups of nodes. One of the most important issues encountered when modeling bipartite networks is devising a way to obtain a (monopartite) projection on the layer of interest, which preserves as much as possible the information encoded into the original bipartite structure. In the present paper we propose an algorithm to obtain statistically-validated projections of bipartite networks, according to which any two nodes sharing a statistically-significant number of neighbors are linked. Since assessing the statistical significance of nodes similarity requires a proper statistical benchmark, here we consider a set of four null models, defined within the exponential random graph framework. Our algorithm outputs a matrix of link-specific p-values, from which a validated projection is straightforwardly obtainable, upon running a multiple hypothesis testing procedure. Finally, we test our method on an economic network (i.e. the countries-products World Trade Web representation) and a social network (i.e. MovieLens, collecting the users' ratings of a list of movies). In both cases non-trivial communities are detected: while projecting the World Trade Web on the countries layer reveals modules of similarly-industrialized nations, projecting it on the products layer allows communities characterized by an increasing level of complexity to be detected; in the second case, projecting MovieLens on the films layer allows clusters of movies whose affinity cannot be fully accounted for by genre similarity to be individuated.Comment: 16 pages, 9 figure