On the dynamics of multi-species Ricker models admitting a carrying simplex

We study the dynamics of the Ricker model (map) T. It is known that under mild conditions, T admits a carrying simplex , which is a globally attracting invariant hypersurface of codimension one. We define an equivalence relation relative to local stability of fixed points on the boundary of Σ on the space of all 3D Ricker models admitting carrying simplices. There are a total of 33 stable equivalence classes. We list them in terms of simple inequalities on the parameters, and draw each one's phase portrait on Σ. Classes 1-18 have trivial dynamics, i.e. every orbit converges to some fixed point. Each map from classes 19-25 admits a unique positive fixed point with index -1, and Neimark-Sacker bifurcations do not occur in these 7 classes. In classes 26-33, there exists a unique positive fixed point with index 1. Within each of classes 26 to 31, there do exist Neimark-Sacker bifurcations, while in class 32 Neimark-Sacker bifurcations can not occur. Whether there is a Neimark-Sacker bifurcation in class 33 or not is still an open problem. Class 29 can admit Chenciner bifurcations, so two isolated closed invariant curves can coexist on the carrying simplex in this class. Each map in class 27 admits a heteroclinic cycle, i.e. a cyclic arrangement of saddle fixed points and heteroclinic connections. As the growth rate increases the carrying simplex will break, and chaos can occur for large growth rate. We also numerically show that the 4D Ricker map can admit a carrying simplex containing a chaotic attractor, which is found in competitive mappings for the first time.Peer reviewe

A survey on the analysis and control of evolutionary matrix games

In support of the growing interest in how to efficiently influence complex systems of interacting self interested agents, we present this review of fundamental concepts, emerging research, and open problems related to the analysis and control of evolutionary matrix games, with particular emphasis on applications in social, economic, and biological networks. (C) 2018 Elsevier Ltd. All rights reserved

PERMANENCE AND UNIVERSAL CLASSIFICATION OF DISCRETE-TIME COMPETITIVE SYSTEMS VIA THE CARRYING SIMPLEX

We study the permanence and impermanence for discrete-time Kolmogorov systems admitting a carrying simplex. Sufficient conditions to guarantee permanence and impermanence are provided based on the existence of a carrying simplex. Particularly, for low-dimensional systems, permanence and impermanence can be determined by boundary fixed points. For a class of competitive systems whose fixed points are determined by linear equations, there always exists a carrying simplex. We provide a universal classification via the equivalence relation relative to local dynamics of boundary fixed points for the three-dimensional systems by the index formula on the carrying simplex. There are a total of 33 stable equivalence classes which are described in terms of inequalities on parameters, and we present the phase portraits on their carrying simplices. Moreover, every orbit converges to some fixed point in classes 1-25 and 33; there is always a heteroclinic cycle in class 27; Neimark-Sacker bifurcations may occur in classes 26-31 but cannot occur in class 32. Based on our permanence criteria and the equivalence classification, we obtain the specific conditions on parameters for permanence and impermanence. Only systems in classes 29,31,33 and those in class 27 with a repelling heteroclinic cycle are permanent. Applications to discrete population models including the Leslie-Gower models, Atkinson-Allen models and Ricker models are given.Peer reviewe

Nonlinear oscillation and control in the BZ chemical reaction.

In this thesis, a reversible Lotka-Volterra model was proposed to study the nonlinear oscillation of the Belousov-Zhabotinsky(BZ) reaction in a closed isothermal chemical system. The reaction zone can be divided into two zones, oscillation zone and transition zone, where the oscillation time and the transition time and the number of the complete oscillations are estimated. By applying the geometric singular perturbation method, it was proved that there exist an oscillation axis in the oscillation zone, a strongly stable two-dimensional invariant manifold in transition zone, on which there is also a one-dimensional stable invariant manifold, which is the part of the central axis. There is no oscillation in the vicinity of the equilibrium, as indicated by Onsager's reciprocal symmetry relation. Furthermore, the damped oscillation is studied in terms of the action-action-angle variables. In the end, the model reference control technique is employed to control the oscillation amplitude in the BZ reaction.Ph.D.Committee Chair: Yi, Yingfei; Committee Member: Chow, Shui-Nee; Committee Member: Dieci, Luca; Committee Member: Verriest, Erik; Committee Member: Weiss, Howi

Balance manifolds in Lotka-Volterra systems

The Lotka-Volterra equations are a dynamical system in the form of an autonomous ODE. The aim of this thesis is to explore the carrying simplex for non-competitive Lotka-Volterra systems for the case of 2- and 3-species, where it is referred to as a balance simplex. Carrying simplices were developed by M.W. Hirsch in a series of papers. They are hypersurfaces which asymptotically attract all non-zero solutions in the phase portrait. This essentially means that all the non-trivial dynamics occur on the carrying simplex, which is one dimension less than the system itself. Many of its properties have been studied by various authors, for example: E.C. Zeeman, M.L. Zeeman, S. Baigent, J. Mierczyński. The first few chapters of this thesis explores the 2-species scaled Lotka-Volterra system, where all intrinsic growth rates and intraspecific interaction rates are set to the value 1. This simplification of the model allows for an explicit, analytic form of the balance simplex to be found. This is done by transforming the system to polar co-ordinates and explicitly integrating the new system. The balance simplex for this 2-species model is precisely composed of the heteroclinic orbits connecting non-zero steady states, along with these states themselves. The later chapters of this thesis focuses on the 3-species case. The existence of the balance simplex in particular parameter cases is proven and it is shown to be piecewise analytic (when the interaction matrix containing the parameters is strictly copositive). These chapters also work towards plotting the balance simplex so it can be visualised for the 3-species system. In another chapter, more general planar Kolmogorov models are considered. Conditions sufficient for the balance simplex to exist are given, and it is again composed of heteroclinic orbits between non-zero steady states

Chaos to Permanence - Through Control Theory

Work by Cushing et al. [18] and Kot et al. [60] demonstrate that chaotic behavior does occur in biological systems. We demonstrate that chaotic behavior can enable the survival/thriving of the species involved in a system. We adopt the concepts of persistence/permanence as measures of survival/thriving of the species [35]. We utilize present chaotic behavior and a control algorithm based on [66, 72] to push a non-permanent system into permanence. The algorithm uses the chaotic orbits present in the system to obtain the desired state. We apply the algorithm to a Lotka-Volterra type two-prey, one-predator model from [30], a ratio-dependent one-prey, two-predator model from [35] and a simple prey-specialist predator-generalist predator (for ex: plant-insect pest-spider) interaction model [67] and demonstrate its effectiveness in taking advantage of chaotic behavior to achieve a desirable state for all species involved

Chaos to Permanence-Through Control Theory

Work by Cushing et al. \cite{Cushing} and Kot et al. \cite{Kot} demonstrate that chaotic behavior does occur in biological systems. We demonstrate that chaotic behavior can enable the survival/thriving of the species involved in a system. We adopt the concepts of persistence/permanence as measures of survival/thriving of the species \cite{EVG}. We utilize present chaotic behavior and a control algorithm based on \cite{Vincent97,Vincent2001} to push a non-permanent system into permanence. The algorithm uses the chaotic orbits present in the system to obtain the desired state. We apply the algorithm to a Lotka-Volterra type two-prey, one-predator model from \cite{Harvesting}, a ratio-dependent one-prey, two-predator model from \cite{EVG} and a simple prey-specialist predator-generalist predator (for ex: plant-insect pest-spider) interaction model \cite{Upad} and demonstrate its effectiveness in taking advantage of chaotic behavior to achieve a desirable state for all species involved